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Skipper leads the penguin quartet that works like an elite military unit. With Kowalski, Rico and Private, he hijacks ships, breaks blockades and lives on escape plans, usually with more confidence than common sense. In the penguins' own film, he and his crew finally take the lead.
Skipper and the penguins have long planned to escape the zoo. When the zoo decides to ship the escaped animals to Kenya, they hijack the ship and set course for Antarctica. The trip accidentally knocks Alex's crate overboard. In the end they find Antarctica inhospitable and land at Madagascar.
Skipper, Kowalski, Rico and Private fix up a crumbling plane with help from chimps recruited by Mason and Phil. To get parts, they steal tourists' safari vehicles and strip them down. The repaired plane flies again at the end, to help rescue Alex and Zuba.
The penguins and chimps head to Monte Carlo for a honeymoon. Skipper and the crew win a fortune at the casino and use it to buy Circus Zaragoza from its human ringmaster. Alex's gang finds them there, and the two groups end up in the same circus.
In the penguins' film, Skipper, Rico and Kowalski rescue an egg in Antarctica, and out comes Private, adopted as a brother. Ten years later, the quartet faces Dave, a vengeful octopus, and the North Wind, who distrust them. In the end, Skipper declares Private the team's most valued member.
Tom McGrath, co-director of the first two films, voices Skipper. The penguins began in a different project, Rockumentary, and later got the Nickelodeon series The Penguins of Madagascar (2008–2015) and the 2014 film.
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